Top100 animals

Saturday, April 7, 2007

Number 93 Tropical Frogs

The frog is an amphibian in the order Anura (meaning "tail-less" from Greek an-, without + oura, tail), formerly referred to as Salientia (Latin saltare, to jump). Adult frogs are characterised by long hind legs, a short body, webbed digits, protruding eyes and the absence of a tail. Most frogs have a semi-aquatic lifestyle, but move easily on land by jumping or climbing. They typically lay their eggs in puddles, ponds or lakes; and their larvae, called tadpoles, have gills and develop in water. Adult frogs follow a carnivorous diet, mostly of arthropods, annelids and gastropods. Frogs are most noticeable by their call, which can be widely heard during the night or day, mainly in their mating season.
The distribution of frogs ranges from tropic to subarctic regions, but most species are found in tropical rainforests. Consisting of more than 5,000 species described, they are among the most diverse groups of vertebrates. However, populations of certain frog species are significantly declining.
A distinction is often made between frogs and toads on the basis of their appearance, caused by the convergent adaptation among so-called toads to dry environments; however, this distinction has no taxonomic basis. The only family exclusively given the common name "toad" is Bufonidae, but many species from other families are also called "toads," and the species within the toad genus Atelopus are referred to as "harlequin frogs."
The name frog derives from Old English frocga,[1] which in turn comes from Old Norse frauki, cognate with Sanskrit plava (frog), probably deriving from Proto-Indo-European praw (to jump).[2] The scientific study of frogs is called herpetology (Greek ἑρπετόν herpeton = to creep, to ramp and λόγος logos = in this context, explanation or reason), a field in zoology that includes the study of both reptiles and amphibians.The life cycle of a frog starts with an egg. A female generally lays frogspawn, or egg masses containing thousands of eggs, in water. The eggs are highly vulnerable to predation, so frogs have evolved many techniques to ensure the survival of the next generation. Most commonly, this involves synchronous reproduction. Many individuals will breed at the same time, overwhelming the actions of predators; the majority of the offspring will still die due to predation, but there is a greater chance that some will survive. Another way in which some species avoid the predators and pathogens eggs are exposed to in ponds is to lay eggs on leaves above the pond, with a gelatinous coating designed to retain moisture. In these species the tadpoles drop into the water upon hatching. The eggs of some species laid out of water can detect vibrations of nearby predatory wasps or snakes, and will hatch early to avoid being eaten.[14] Some species, such as the Cane Toad (Bufo marinus), lay poisonous eggs to minimise predation. While the length of the egg stage depends on the species and environmental conditions, aquatic eggs generally hatch within one week.
Eggs hatch and continue life as tadpoles (occasionally known as polliwogs). Tadpoles are aquatic, lack front and hind legs, and have gills for respiration and tails with fins for swimming. Tadpoles are typically herbivorous, feeding mostly on algae, including diatoms that are filtered from the water through the gills. Some species are carnivorous at the tadpole stage, eating insects, smaller tadpoles and fish. Tadpoles are highly vulnerable to predation by fish, newts, predatory diving beetles and birds such as kingfishers. Cannibalism has been observed among tadpoles. Poisonous tadpoles are present in many species, such as Cane Toads. The tadpole stage may be as short as a week, or tadpoles may overwinter and metamorphose the following year in some species, such as the Midwife toad (Alytes obstetricans) and the Common Spadefoot (Pelobates fuscus).
At the end of the tadpole stage, frogs undergo metamorphosis, in which they transition into adult form. Metamorphosis involves a dramatic transformation of morphology and physiology, as tadpoles develop hind legs, then front legs, lose their gills and develop lungs. Their intestines shorten as they shift from an herbivorous to a carnivorous diet. Eyes migrate rostrally and dorsally, allowing for binocular vision exhibited by the adult frog. This shift in eye position mirrors the shift from prey to predator, as the tadpole develops and depends less upon a larger and wider field of vision and more upon depth perception. The final stage of development from froglet to adult frog involves apoptosis (programmed cell death) and resorption of the tail.
After metamorphosis, young adults may leave the water and disperse into terrestrial habitats, or continue to live in the aquatic habitat as adults. Almost all species of frogs are carnivorous as adults, eating invertebrates such as arthropods, annelids and gastropods. A few of the larger species may eat prey such as small mammals, fish and smaller frogs. Some frogs use their sticky tongues to catch fast-moving prey, while others capture their prey and force it into their mouths with their hands. However, there are a very few species of frogs that primarily eat plants.[15] Adult frogs are themselves preyed upon by birds, large fish, snakes, otters, foxes, badgers, coatis, and other animals. Frogs are also eaten by people (see section on agriculture, below).

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